Taxa modified
Nectaspis areolate renamed
Nectaspis areolata
Vernicomacanthus waynensis renamed Dobunnacanthus
waynensis
Taxa added
Ateleaspis tessellata
Kolymaspis
sibirica
Minjinia
turgenensis
Characters
deleted (numberings from King et al. 2017):
Character
22 (Elongate preorbital region between orbits and nasal capsules) deleted.
Character
173 (ZHU169, GFB60 Type of dermal neck-joint) deleted.
Character
174 (Type of ginglymoid neck joint) deleted.
Character
192 (ZHU168 Dermal neck-joint between paired main-lateral-line-bearing bones of
skull) deleted.
Characters
added:
Character 7
(Laminar basal layer) added.
Character 8
(Cancellous central layer) added.
Character 9
(Dentinous external layer) added.
Character
25 (Extended preorbital region between eyes and nasal capsule) added.
Character
52 (Crus commune) added.
Character
177 (ZHU168* Dermal cranio-thoracic articulation between paired
main-lateral-line-bea) added.
Character
178 (Dermal neck joint facet morphology) added.
Character
179 (Para-articular process of dermal neck joint) added.
Character
180 (Endoskeletal craniothoracic (sixth branchial)
facet) added.
Table of state score changes
Taxon |
Orig. char. no. |
New char. no. |
Character label |
Old state |
New state |
Justification |
Acanthodes bronni |
140 |
144 |
(DFC37 Basihyal) |
1 (absent, hyoid arch
articulates directly with basibranchial) |
0 (present) |
We code Acanthodes as having a basihyal
after Dearden et al. (2019) |
Achoania jarvikii |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Achoania jarvikii |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Achoania jarvikii |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Benneviaspis holtedahli |
96 |
100 |
(Nasal capsules in anterlolateral corners of orbit) |
0 (no) |
- (inapplic.) |
Although olfactory bulbs
are present, discrete nasal capsules appear to be absent in osteostracans. |
Benneviaspis holtedahli |
422 |
427 |
(DFC105 Dermal shoulder
girdle composition) |
0 (ventral and dorsal
(scapular) components) |
- (inapplic.) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |
Boreaspis macrorhynchus |
96 |
100 |
(Nasal capsules in anterlolateral corners of orbit) |
0 (no) |
- (inapplic.) |
Although olfactory bulbs
are present, discrete nasal capsules appear to be absent in osteostracans. |
Boreaspis macrorhynchus |
422 |
427 |
(DFC105 Dermal shoulder
girdle composition) |
0 (ventral and dorsal (scapular)
components) |
- (inapplic.) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |
Brindabellaspis stensioi |
48 |
51 |
(DFC86 Sinus superior) |
0 (absent or
indistinguishable from union of anterior and posterior canals with saccular
chamber) |
1 (present) |
Updated based on Zhu et
al. (2021, Curr. Biol.) |
Brindabellaspis stensioi |
80 |
84 |
([GFB141]Subcranial ridges) |
- (inapplic.) |
1 (present) |
We consider the laterobasal angles of brindabellaspidids
to be primary homologues of these ridges based on topological position. |
Brindabellaspis stensioi |
97 |
101 |
(vagal
process) |
- (inapplic.) |
0 (forked) |
A vagal process (or its
homologue) is evidently present as documented by Young (1989. Palaeontographica A, figs. 7, 13) |
Brindabellaspis stensioi |
114 |
118 |
(Hypophysial foramen in
braincase) |
1 (present) |
0 (absent) |
There is no evidence of
a foramen according to Young (1980) and Zhu et al. (2021) |
Brindabellaspis stensioi |
187 |
193 |
(ZHU159 Parietals
(preorbitals of placoderms) surround pineal foramentotoeminence) |
1 (no) |
0 (yes) |
Updated based on Zhu et
al. (2021, Curr. Biol.) |
Brindabellaspis stensioi |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Brindabellaspis stensioi |
248 |
252 |
(Preorbital recess) |
- (inapplic.) |
0 (absent) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Cephalaspis lyelli |
422 |
427 |
(DFC105 Dermal shoulder
girdle composition) |
0 (ventral and dorsal
(scapular) components) |
- (inapplic.) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |
Cephalaspis lyelli |
423 |
428 |
(DFC106 Dermal shoulder
girdle forming a complete ring around the trunk) |
0 (present) |
1 (absent) |
|
Cheirolepis canadensis |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Cheirolepis canadensis |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Cheirolepis canadensis |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Cheirolepis trailli |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Cheirolepis trailli |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Cheirolepis trailli |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Diabolepis speratus |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Diabolepis speratus |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Diabolepis speratus |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Entelognathus primordialis |
84 |
88 |
([GFB166]Branchial
ridges) |
2 (absent (articulation
made with bare cranial wall)) |
1 (reduced to vagal
process) |
Zhu et al. (2013) figure
a small posterior postorbital process in Entelognathus
which is homologizeable with the vagal process
(after Giles et al. 2015, Nature). |
Escuminaspis laticeps |
422 |
427 |
(DFC105 Dermal shoulder
girdle composition) |
0 (ventral and dorsal
(scapular) components) |
- (inapplic.) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |
Eusthenopteron foordi |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Eusthenopteron foordi |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Eusthenopteron foordi |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Gemuendina stuertzi |
186 |
192 |
(ZHU158 Location of
pineal foramen/eminence) |
1 (well posterior of
orbits) |
0 (level with posterior
margin of orbits) |
The pineal opening of rhenanids is situated between the orbits and therefore
cannot be considered well posterior to them (Gross 1963; Stensiö,
1969) |
Gemuendina stuertzi |
246 |
251 |
(Preorbital depression) |
- (inapplic.) |
1 (present) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Glyptolepis groenlandica |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Glyptolepis groenlandica |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Glyptolepis groenlandica |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Gogonasus andrewsae |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Gogonasus andrewsae |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Gogonasus andrewsae |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Guiyu oneiros |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Guiyu oneiros |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Guiyu oneiros |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Hemicyclaspis murchisoni |
422 |
427 |
(DFC105 Dermal shoulder
girdle composition) |
0 (ventral and dorsal
(scapular) components) |
- (inapplic.) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |
Hemicyclaspis murchisoni |
423 |
428 |
(DFC106 Dermal shoulder
girdle forming a complete ring around the trunk) |
0 (present) |
1 (absent) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |
Howqualepis rostridens |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Howqualepis rostridens |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Howqualepis rostridens |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Jagorina pandora |
186 |
192 |
(ZHU158 Location of pineal
foramen/eminence) |
1 (well posterior of
orbits) |
0 (level with posterior
margin of orbits) |
The pineal opening of rhenanids is situated between the orbits and therefore
cannot be considered well posterior to them (Gross 1963; Stensiö,
1969) |
Kenichthys campbelli |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Kenichthys campbelli |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Kenichthys campbelli |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Kentuckia deani |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Kentuckia deani |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Kentuckia deani |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Lunaspis broili |
158 |
162 |
(DFC25 Pineal opening
perforation in dermal skull roof) |
1 (absent) |
0 (present) |
A pineal opening is
present in Lunaspis (Gross 1961, fig. 1) |
Lunaspis broili |
450 |
455 |
(DFC136 Number of dorsal
fins, if present) |
0 (one) |
- (inapplic.) |
Dorsal fins are absent
in Lunaspis (Gross 1961) |
Microbrachius dicki |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Microbrachius dicki |
248 |
252 |
(Preorbital recess) |
- (inapplic.) |
0 (absent) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Mimipiscis toombsi |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Mimipiscis toombsi |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Mimipiscis toombsi |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Moythomasia durgaringa |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Moythomasia durgaringa |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Moythomasia durgaringa |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Norselaspis glacialis |
96 |
100 |
(Nasal capsules in anterlolateral corners of orbit) |
0 (no) |
- (inapplic.) |
Although olfactory bulbs
are present, discrete nasal capsules appear to be absent in osteostracans. |
Norselaspis glacialis |
422 |
427 |
(DFC105 Dermal shoulder
girdle composition) |
0 (ventral and dorsal
(scapular) components) |
- (inapplic.) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |
Norselaspis glacialis |
423 |
428 |
(DFC106 Dermal shoulder
girdle forming a complete ring around the trunk) |
0 (present) |
1 (absent) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |
Onychodus jandemarrai |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Onychodus jandemarrai |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Onychodus jandemarrai |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Osorioichthys marginis |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Osorioichthys marginis |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Osorioichthys marginis |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Osteolepis macrolepidotus |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Osteolepis macrolepidotus |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Osteolepis macrolepidotus |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Porolepis sp |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Porolepis sp |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Porolepis sp |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Powichthys thorsteinssoni |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Powichthys thorsteinssoni |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Powichthys thorsteinssoni |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Procephalaspis oeselensis |
96 |
100 |
(Nasal capsules in anterlolateral corners of orbit) |
0 (no) |
- (inapplic.) |
Although olfactory bulbs
are present, discrete nasal capsules appear to be absent in osteostracans. |
Procephalaspis oeselensis |
422 |
427 |
(DFC105 Dermal shoulder
girdle composition) |
0 (ventral and dorsal
(scapular) components) |
- (inapplic.) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |
Psarolepis romeri |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Psarolepis romeri |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Psarolepis romeri |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Pterichthyodes milleri |
247 |
253 |
(Preorbital recess) |
1 (present) |
0 (restricted to premedian plate) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Pterichthyodes milleri |
248 |
252 |
(Preorbital recess) |
0 (restricted to premedian plate) |
1 (present) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Remigolepis walkeri |
247 |
253 |
(Preorbital recess) |
1 (present) |
0 (restricted to premedian plate) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Remigolepis walkeri |
248 |
252 |
(Preorbital recess) |
0 (restricted to premedian plate) |
1 (present) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Romundina stellina |
48 |
51 |
(DFC86 Sinus superior) |
1 (present) |
0 (absent or indistinguishable
from union of anterior and posterior canals with saccular chamber) |
Updated based on Dupret et al. (2017, PLOS ONE) |
Romundina stellina |
131 |
135 |
(DFC55 Median dermal
bone of palate (parasphenoid)) |
1 (present) |
0 (absent) |
There is no concrete
evidence of a parasphenoid in Romundina. Dupret et al. (2017) identified some thickened
perichondrium in the hypophyseal region, but stop
short of identifying it as a parasphenoid. |
Romundina stellina |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Romundina stellina |
248 |
252 |
(Preorbital recess) |
- (inapplic.) |
0 (absent) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Shuyu zhejiangensis |
48 |
51 |
(DFC86 Sinus superior) |
0 (absent or
indistinguishable from union of anterior and posterior canals with saccular
chamber) |
1 (present) |
Gai et al. (2011)
supplementary movie shows what is a distinct sinus superior, contra Zhu et
al. (2021. Curr. Biol.) |
Styloichthys changae |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Styloichthys changae |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Styloichthys changae |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Tremataspis mammillata |
96 |
100 |
(Nasal capsules in anterlolateral corners of orbit) |
0 (no) |
- (inapplic.) |
Although olfactory bulbs
are present, discrete nasal capsules appear to be absent in osteostracans. |
Tremataspis mammillata |
422 |
427 |
(DFC105 Dermal shoulder
girdle composition) |
0 (ventral and dorsal
(scapular) components) |
- (inapplic.) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |
Waengsjoeaspis excellens |
422 |
427 |
(DFC105 Dermal shoulder
girdle composition) |
0 (ventral and dorsal
(scapular) components) |
- (inapplic.) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |
Youngolepis praecursor |
246 |
251 |
(Preorbital depression) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Youngolepis praecursor |
247 |
252 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Youngolepis praecursor |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Yunnanolepis sp |
247 |
253 |
(Preorbital recess) |
0 (absent) |
- (inapplic.) |
Character is not applicable
to taxa without an upper lip/premedian area. |
Yunnanolepis sp |
248 |
252 |
(Preorbital recess) |
- (inapplic.) |
0 (absent) |
Character is not
applicable to taxa without an upper lip/premedian
area. |
Zenaspis salweyi |
422 |
427 |
(DFC105 Dermal shoulder
girdle composition) |
0 (ventral and dorsal
(scapular) components) |
- (inapplic.) |
We code this character
as inapplicable for osteostracans because our comparative anatomical model
proposes that a dorsal area is not yet present in these taxa (and forms
during a dorsal expansion of the pharynx) |