Taxa, sources formations and age used in analysis

 

Original source: King B., Qiao T., Lee M.S.Y., Zhu M., Long J.A. 2017. Bayesian morphological clock methods resurrect placoderm monophyly and reveal rapid early evolution in jawed vertebrates. Systematic Biology. 66:499–516.

 

Taxon

Source(s)

Formation

Age for analysis

Ateleaspis tessellata

(Ritchie 1967)

Slot Burn Formation

432

Hemicyclaspis

murchisoni

(Stensiö 1932)

Ludlow, Shropshire.

Downtownian (Pridoli)

421

Zenaspis salweyi

(Stensiö 1932)

Lower old red sandstone. Skirrid Fawr, Senni/St

Maughans formation

412

Cephalaspis lyelli

(Stensiö 1932, White

1958)

Lower old red

sandstone, Glammis

412

Benneviaspis

holtedahli

(Janvier 1985a)

Ben Nevis formation,

Red bay group.

413

Boreaspis

macrorhynchus

(Janvier 1985a)

Wood bay formation

411

Norselaspis glacialis

(Janvier 1981)

Wood bay formation

411

Nectaspis areolata

(Wängsjö 1952, Janvier

1981)

Wood bay formation

411

Procephalaspis oeselensis

(Robertson 1939,

Denison 1951, Janvier 1985b)

Saaremaa, Estonia. Wenlockian and

Ludlowian

427

Tremataspis mammillata

(Robertson 1938a, Robertson 1938b, Denison 1947, Denison

1951, Janvier 1985b)

Saaremaa

427

Waengsjoeaspis

excellens

(Wängsjö 1952, Janvier

1985a)

Fraenkelryggen

formation

416

Escuminaspis laticeps

(Janvier et al. 2004)

Escuminac formation

380

Eugaleaspis changi

(Liu 1965, Zhu and Gai

2007)

Xitun formation,

Liaojaoshan

412

Hanyangaspis

guodingshanensis

(Zhu and Gai 2007)

Guodingshan

formation. Telychian

436

Polybranchiaspis

liaojiaoshanensis

(Liu 1965, Liu 1975)

Xishancun and Xitun

formations

415

Bannhuanaspis

vukhuci

(Janvier et al. 1993)

Bac Bun formation

411

Wenshanaspis zhichangensis

(Zhao et al. 2001)

Posongchong

formation, Wenshan. Pragian

409

Shuyu zhejiangensis

(Gai et al. 2011)

Maoshan formation. Late Telychian to early

Wenlock

433

Polybranchiaspid sp.

(histological samples)

(Wang et al. 2005)

Xishancun and Xitun

formations

415

Yunnanolepis sp. (various taxa from

same formations)

(Zhang 1980, Zhu 1996)

Xishancun and Xitun formations

415

Parayunnanolepis

xitunensis

(Zhang et al. 2001, Zhu

et al. 2012)

Xitun formation

412

Microbrachius dicki

(Hemmings 1978, Long

et al. 2015)

Eday flagstone and John O’Groats sandstone. Lower-

Middle Givetian

386

Bothriolepis sp.

(Young 1984)

MVP230985

Gogo formation

383

Bothriolepis canadensis

(Downs and Donoghue 2009, Béchard et al.

2014)

Escuminac formation

380

Pterichthyodes milleri

(Hemmings 1978)

Achanarras horizon, under upper Stromness flagst. Late

Eifelian

389

Remigolepis walkeri

(Johanson 1997)

Canowindra.

Famennian

366

Diandongpetalichthys

liaojiaoshanensis

(Zhu 1991)

Xishancun formation

417

Quasipetalichthys haikouensis

(Liu 1991)

Kunming; Shixiagou formation, Ninxia.

Givetian

385

Eurycaraspis incilis

(Liu 1991)

Haikou formation.

Givetian

385

Lunaspis broili

(Gross 1961)

Hunsrueck slate. Late Pragian to early

Emsian

408

Macropetalichthys rapheidolabis. (including Macropetalichthys sp. specimens used in

1963 and 1969)

(Stensiö 1925, Stensiö

1963b, Stensiö 1969)

Onondaga limestone. Eifelian

390

Wuttagoonaspis fletcheri

(Ritchie 1973, Miles and Young 1977).

AMF53610, AMF53631,

AMF53591,

Mulga Downs group. Emsian-Eifelian

393

Groenlandaspis sp.

MVP48873(AMF59808) , AMF59814, MVP48877/8 (AMF59811/2),

AMF62534A, AMF59809

Mt. Howitt

385

Cowralepis mclachlani

(Ritchie 2005, Carr et al. 2009)

Merriganowry shale. Late Givetian to early

Frasnian

383

Gavinaspis convergens

(Dupret et al. 2009)

Xitun formation.

412

Sigaspis lepidophora

(Goujet 1973)

Wood bay formation

(extreme base)

412

Kujdanowiaspis podolica

(Stensiö 1963a, Dupret 2010)

Dnister series, Podolia. Upper Lockhovian-

lower Pragian

411

Dicksonosteus arcticus

(Goujet 1975, Goujet

1984)

Wood bay formation

411

Buchanosteus

confertituberculatus

(Burrow and Turner

1998, Long et al. 2014)

Buchan. Mid-late

Pragian

408

Parabuchanosteus murrumbidgeensis

(White and Toombs 1972, Young 1979,

Burrow and Turner

1998)

Taemas-Wee Jasper

401

Holonema westolli

(Miles 1971)

Gogo formation

383

Coccosteus cuspidatus

(Miles and Westoll 1968)

Achanarras and Edderton fish bed. Eifelian-Givetain

boundary

388

Incisoscutum ritchiei

(Dennis and Miles 1981,

Giles et al. 2013)

Gogo formation

383

Eastmanosteus

calliaspis

(Dennis-Bryan 1987)

Gogo formation

383

Compagopiscis

croucheri

(Gardiner and Miles

1994)

Gogo formation

383

Materpiscis

attenboroughi

(Long et al. 2008,

Trinajstic et al. 2012)

Gogo formation

383

Austroptyctodus

gardineri

(Long 1997a)

Gogo Formation

383

Campbellodus

decipiens

(Long 1997a)

Gogo formation

383

Rhamphodopsis threiplandi

(Miles 1967, Long 1997b)

Edderton fish beds. Eifelian-Givetian

boundary

388

Brindabellaspis

stensioi

(Young 1980).

ANUV3247

Taemas-Wee Jasper

401

Kolymaspis sibirica

TsNIGR 7656

Vechernyaya Formation

400

Romundina stellina

(Ørvig 1975, Dupret et

al. 2014)

Lockhovian. Prince of

Wales island.

415

Jagorina pandora

(Stensiö 1969, Young

1986)

Kellwasserkalk, Bad

Wildungen. Late Frasnian

375

Gemuendina stuertzi

(Gross 1963)

Hunsrueck slate

408

Minjinia turgenensis

(Brazeau et al. 2020)

Tsagaansalaa Formation (Baastuul Formation)

410

Entelognathus

primordialis

(Zhu et al. 2013)

Kuanti formation

424

Janusiscus schultzei

(Giles et al. 2015b)

Lower member, Kureika formation.

Middle Lockhovian.

415

Ramirosuarezia

boliviana

(Pradel et al. 2009)

Icla formation. Early

Eifelian

392

Acanthodes bronni

(Gross 1935, Watson

1937, Miles 1973b, Miles 1973a, Coates 1994, Davis et al. 2012, Brazeau and de Winter

2015)

Lebach ironstone. Early Permian

298

Brachyacanthus scutiger

(Watson 1937)

Lower old red sandstone, Farnell.

Lockhovian

415

Brochoadmones milesi

(Hanke and Wilson

2006)

MOTH

415

Cassidiceps

vermiculatus

(Gagnier and Wilson

1996)

MOTH

415

Cheiracanthus latus/murchisoni

(Watson 1937, Miles 1973a)

Middle old red sandstone, Moray firth. Nodular fish

beds, Eifelian-Givetian

388

Climatius reticulatus

(Watson 1937, Miles

1973a)

‘Turin hill’

415

Culmacanthus stewarti

(Long 1983)

Mt Howitt

385

Euthacanthus macnicoli

(Watson 1937, Miles 1973a, Newman et al.

2014)

‘Turin hill’

415

Gladiobranchus

probation

(Hanke and Davis 2008)

MOTH

415

Homalacanthus

concinnus

(Gagnier 1996)

Escuminac formation

380

Ischnacanthus gracilis

(Watson 1937, Miles

1973a)

‘Turin hill’

415

Kathemacanthus rosulentus

(Gagnier and Wilson

1996, Hanke and Wilson 2010)

MOTH

415

Latviacanthus

ventspilensis

(Schultze and Zidek

1982)

Ventspils. Kemeri

stage

404

Lupopsyrus pygmaeus

(Hanke and Davis 2012)

MOTH

415

Mesacanthus mitchelli

(Watson 1937, Miles

1973a)

‘Turin hill’ and Farnell

415

Obtusacanthus

corroconis

(Hanke and Wilson

2004)

MOTH

415

Parexus recurvus

(Watson 1937, Miles 1973a, Burrow et al.

2013)

‘Turin hill’

415

Poracanthodes menneri

(Valiukevicius 1992)

Severnaya Zemlya

formation. Lower Lockhovian.

417

Promesacanthus

eppleri

(Hanke 2008)

MOTH

415

Ptomacanthus anglicus

(Miles 1973a, Brazeau

2012)

Wayne Herbert

quarry, Lockhovian

415

Diplacanthus striatus

(Watson 1937, Miles 1973a)

Moray firth and Achanarras. Eifelian-

Givetian boundary

388

Tetanopsyrus

lindoei/breviacanthus

(Gagnier et al. 1999,

Hanke et al. 2001)

MOTH

415

Dobunnacanthus

waynensis

(Miles 1973a, Dearden et al. 2021)

Wayne Herbert quarry

415

Cladodoides

wildungensis

(Maisey 2005)

Wildungen limestone,

Upper Frasnian

375

Akmonistion zangerli

(Coates and Sequeira 1998, Coates et al.

1998, Coates and

Sequeira 2001)

Manse Burn formation, Bearsden. Serpukhovian

327

Cobelodus

(Maisey 2007)

Fayetteville formation.

Chesterian

325

Cladoselache kepleri/fyerli

(Harris 1938, Bendix-

Almgreen 1975,

Schaeffer 1981, Maisey

2007)

Cleveland member of Ohio Shale. Late Famennian

360

Chondrenchelys problematica

(Moy-Thomas 1935, Finarelli and Coates 2011, Finarelli and

Coates 2014)

Glencartholm, Scotland. Visean (Holkerian)

338

Helodus simplex

(Moy-Thomas 1936)

Fenton, Staffordshire.

Moscovian

311

Debeerius ellefseni

(Grogan and Lund 2000)

Bear Gulch limestone.

Upper Chesterian

320

Doliodus problematicus

(Miller et al. 2003, Maisey et al. 2009)

Atholville beds’, Campbellton formation. Emsian or

Emsian-Eifelian

395

Hamiltonichthys mapesi

(Maisey 1989)

Hartford limetone, Hamilton quarry.

Middle Virgilian

302

Onychoselache traquari

(Dick and Maisey 1980, Coates and Gess 2007)

Glencartholm volcanic beds and Wardie shales. Holkerian-

Asbian

336

Orthacanthus sp.

(Schaeffer 1981)

Admiral formation.

Wolfcampian

290

Pucapampella rodrigae/sp.

(Maisey 2001)

Sica sica formation,

Bolivia. Eifelian- Givetian

388

Tamiobatis vetustus

(Schaeffer 1981,

Williams 1998)

Cleveland shale (Famennian) and Salem limestone (Early

Visean)

360

Tristychius arcuatus

(Dick 1978, Coates and Gess 2007)

Wardie shales and Manse burn

formation, Bearsden

336

Dialipina

salgueiroensis

(Schultze 1968, Schultze

and Cumbaa 2001)

Bear Rock formation.

Emsian

401

Ligulalepis toombsi

(Schultze 1968, Burrow 1994, Basden et al.

2000, Basden and Young

2001)

Taemas-Wee Jasper

401

Cheirolepis canadensis

(Pearson and Westoll 1979, Arratia and

Cloutier 1996)

Escuminac formation

380

Cheirolepis trailli

(Pearson and Westoll 1979, Giles et al. 2015a)

Achanarras limestone, Tynet burn and

Gamrie. Late Eifelian

388

Howqualepis

rostridens

(Long 1988)

Mt. Howitt

385

Mimipiscis toombsi

(Gardiner and Bartram 1977, Gardiner 1984, Giles and Friedman

2014)

Gogo formation

383

Moythomasia

durgaringa

(Gardiner 1984)

Gogo formation

383

Kentuckia deani

(Rayner 1952, Giles and Friedman 2014)

New Providence Shale member, Stockdale formation. Latest Tournasian or earliest

Visean

347

Osorioichthys marginis

(Taverne 1997)

Femenne formation.

Early Femennian

367

Meemannia eos

(Zhu et al. 2006, Zhu et

al. 2010)

Xitun formation

412

Guiyu oneiros

(Zhu et al. 2009, Qiao

and Zhu 2010)

Kuanti formation

424

Psarolepis romeri

(Yu 1998, Zhu et al. 1999, Zhu and Yu 2004, Zhu and Yu 2009)

Xishancun, Xitun and Yulongsi formations. Also Silurian. Most material used to score characters is from

Xishancun formation.

416

Achoania jarvikii

(Zhu et al. 2001, Zhu and Yu 2004, Zhu and

Yu 2009)

Xitun formation

412

Onychodus

jandemarrai

(Andrews et al. 2005)

Gogo formation,

Saddler formation

383

Miguashaia bureaui

(Cloutier 1996, Forey

1998)

Escuminac formation

380

Styloichthys changae

(Zhu and Yu 2002,

Friedman 2007)

Xitun formation

412

Diabolepis speratus

(Chang and Yu 1984,

Chang 1995)

Xitun formation

412

Youngolepis praecursor

(Zhang and Yu 1981, Zhang 1982, Chang

1991)

Xitun formation

412

Powichthys thorsteinssoni

(Jessen 1980)

Late lockhovian or early Pragian. Prince

of Wales island

411

Porolepis sp. (includes elongata, brevis and

spitzbergensis)

(Jarvik 1972, Clement

2004)

Wood bay formation

411

Glyptolepis groenlandica

(Jarvik 1972, Ahlberg

1989)

Red siltstone member of the Nathorst Fjord group. Late Eifelian-

Early Givetian

388

Kenichthys campbelli

(Chang and Zhu 1993, Zhu and Ahlberg 2004)

Chuangdong formation. Late

Emsian

396

Osteolepis

macrolepidotus

(Westoll 1936, Thomson

1965, Jarvik 1980)

Tynet burn. Late

Eifelian

388

Gogonasus andrewsi

(Long 1985, Long et al. 1997, Long et al. 2006,

Holland 2013, Holland

2014)

Gogo formation

383

Eusthenopteron foordi

(Jarvik 1980)

Escuminac formation

380


 

Key of prefixes in character list

MDB”=Brazeau, M.D. 2009.

DFC”= Davis et al. 2012.

ZHU”= Zhu et al. 2013.

DUPRET”= Dupret et al. 2014.

LONG”= Long et al. 2015.

GFB”= Giles et al. 2015.

BW”= Brazeau and de Winter 2015.

“LU” =Lu et al. 2016[BMD1] 

RSS”=Sansom 2009.

 

 


References

 

Ahlberg P.E. 1989. Paired fin skeletons and relationships of the fossil group Porolepiformes (Osteichthyes: Sarcopterygii). Zool. J. Linn. Soc. 96:119-166.

 

Andrews M., Long J., Ahlberg P., Barwick R., Campbell K. 2005. The structure of the sarcopterygian Onychodus jandemarrai n. sp. from Gogo, Western Australia: with a functional interpretation of the skeleton. Transactions of the Royal Society of Edinburgh: Earth Sciences 96:197-307.

 

Arratia G., Cloutier R. 1996. Reassessment of the morphology of Cheirolepis canadensis (Actinopterygii). In: Schultze H.-P., Cloutier R., editors. Devonian Fishes and Plants of Miguasha, Quebec, Canada. . Verlag Dr. Friedrich Pfeil, München: p. 165-197.

 

Bapst D.W. 2012. paleotree: an R package for paleontological and phylogenetic analyses of evolution. Methods in Ecology and Evolution 3:803-807.

 

Basden A.M., Young G.C. 2001. A primitive actinopterygian neurocranium from the Early Devonian of southeastern Australia. J. Vertebr. Paleontol. 21:754-766.

 

Basden A.M., Young G.C., Coates M.I., Ritchie A. 2000. The most primitive osteichthyan braincase? Nature 403:185-188.

 

Béchard I., Arsenault F., Cloutier R., Kerr J. 2014. The Devonian placoderm fish Bothriolepis canadensis revisited with three-dimensional digital imagery. Palaeontologia Electronica 17:1-19.

 

Bendix-Almgreen S. 1975. The paired fins and shoulder girdle in Cladoselache, their morphology and phyletic significance. Colloques International CNRS 218:111-123.

 

Brazeau M.D. 2012. A revision of the anatomy of the Early Devonian jawed vertebrate Ptomacanthus anglicus Miles. Palaeontology 55:355-367.

 

Brazeau M.D., de Winter V. 2015. The hyoid arch and braincase anatomy of Acanthodes support chondrichthyan affinity of ‘acanthodians’. Proc. R. Soc. B 282:20152210.

 

Brazeau M.D., Giles S., Dearden R.P., Jerve A., Ariunchimeg Y., Zorig E., Sansom R., Guillerme T., Castiello M. 2020. Endochondral bone in an Early Devonian “placoderm” from Mongolia. Nat Ecol Evol. 4:1477–1484.

 

Burrow C. 1994. Form and function in scales of Ligulalepis toombsi Schultze, a palaeoniscoid from the Early Devonian of Australia. Records of the South Australian Museum 27:175-185.

 

Burrow C., Turner S. 1998. Devonian placoderm scales from Australia. J. Vertebr. Paleontol. 18:677- 695.

 

Burrow C.J., Newman M.J., Davidson R.G., den Blaauwen J.L. 2013. Redescription of Parexus recurvus, an Early Devonian acanthodian from the Midland Valley of Scotland. Alcheringa: An Australasian Journal of Palaeontology 37:392-414.

 

Carr R.K., Hlavin W.J. 2010. Two new species of Dunkleosteus Lehman, 1956, from the Ohio Shale Formation (USA, Famennian) and the Kettle Point Formation (Canada, Upper Devonian), and a cladistic analysis of the Eubrachythoraci (Placodermi, Arthrodira). Zool. J. Linn. Soc. 159:195-222.

 

Carr R.K., Johanson Z., Ritchie A. 2009. The phyllolepid placoderm Cowralepis mclachlani: Insights into the evolution of feeding mechanisms in jawed vertebrates. J. Morphol. 270:775-804.

 

Chang M.-M. 1991. Head exoskeleton and shoulder girdle of Youngolepis. In: Chang M.-M., Liu Y.-H., Zhang G.-R., editors. Early Vertebrates and Related Problems of Evolutionary Biology. Science Press, Beijing: p. 355-378.

 

Chang M.-M. 1995. Diabolepis and its bearing on the relationships between porolepiforms and dipnoans. Bulletin du Muséum national d'histoire naturelle. Section C, Sciences de la terre, paléontologie, géologie, minéralogie 17:235-268.

 

Chang M.-M., Yu X.-B. 1984. Structure and phlyogenetic significance of Diabolepis speratus gen. et sp. nov., a new dipnoan-like form from the Lower Devonian of eastern Yunnan, China. Proc. Linn. Soc.

NSW, p. 171-184.

 

Chang M.-M., Zhu M. 1993. A new Middle Devonian osteolepidid from Qujing, Yunnan. Memoirs of the Association of Australasian Palaeontologists 15:183-198.

 

Clement G. 2004. Nouvelles données anatomiques et morphologie génerale des Porolepididae (Dipnomorpha, Sarcopterygii). Revue de Paléontologie 9:193-211.

 

Cloutier R. 1996. The primitive actinistian Miguashaia bureaui Schultze (Sarcopterygii). In: Schultze H.-P., Cloutier R., editors. Devonian Fishes and Plants of Miguasha, Quebec, Canada. Verlag Dr. Friedrich Pfeil, München: p. 227-247.

 

Coates M., Sequeira S. 1998. The braincase of a primitive shark. Transactions of the Royal Society of Edinburgh: Earth Sciences 89:63-85.

 

Coates M., Sequeira S. 2001. A new stethacanthid chondrichthyan from the Lower Carboniferous of Bearsden, Scotland. J. Vertebr. Paleontol. 21:438-459.

 

Coates M., Sequeira S., Sansom I., Smith M. 1998. Spines and tissues of ancient sharks. Nature 396:729-730.

 

Coates M.I. 1994. The origin of vertebrate limbs. Development 1994:169-180.

 

Coates M.I. 1999. Endocranial preservation of a Carboniferous actinopterygian from Lancashire, UK, and the interrelationships of primitive actinopterygians. Philosophical Transactions of the Royal Society B: Biological Sciences 354:435-462.

 

Coates M.I., Gess R.W. 2007. A new reconstruction of Onychoselache traquairi, comments on early chondrichthyan pectoral girdles and hybodontiform phylogeny. Palaeontology 50:1421-1446.

 

Davis S.P., Finarelli J.A., Coates M.I. 2012. Acanthodes and shark-like conditions in the last common ancestor of modern gnathostomes. Nature 486:247-250.

 

Dearden R.P., Blaauwen J.L., Sansom I.J., Burrow C.J., Davidson R.G., Newman M.J., Ko A., Brazeau M.D. 2021. A revision of Vernicomacanthus Miles with comments on the characters of stem‐group chondrichthyans. Pap Palaeontol. 7:1949–1976.

 

Denison R.H. 1947. The exoskeleton of Tremataspis. American Journal of Science 245:337-365.

 

Denison R.H. 1951. Evolution and classification of the Osteostraci. Fieldania, Geology 11:157-196. Dennis-Bryan K. 1987. A new species of eastmanosteid arthrodire (Pisces: Placodermi) from Gogo, Western Australia. Zool. J. Linn. Soc. 90:1-64.

 

Dennis K., Miles R. 1981. A pachyosteomorph arthrodire from Gogo, Western Australia. Zool. J. Linn. Soc. 73:213-258.

 

Dick J., Maisey J. 1980. The Scottish Lower Carboniferous shark Onychoselache traquairi. Palaeontology 23:363-374.

 

Dick J.R. 1978. On the Carboniferous shark Tristychius arcuatus Agassiz from Scotland. Trans. R. Soc. Edinb. 70:63-108.

 

Donoghue P.C., Forey P.L., Aldridge R.J. 2000. Conodont affinity and chordate phylogeny. Biological Reviews 75:191-251.

 

Downs J.P., Donoghue P.C. 2009. Skeletal histology of Bothriolepis canadensis (Placodermi, Antiarchi) and evolution of the skeleton at the origin of jawed vertebrates. J. Morphol. 270:1364-1380.

 

Dupret V. 2010. Revision of the genus Kujdanowiaspis Stensiö, 1942 (Placodermi, Arthrodira,“Actinolepida”) from the Lower Devonian of Podolia (Ukraine). Geodiversitas 32:5-63. Dupret V., Sanchez S., Goujet D., Tafforeau P., Ahlberg P.E. 2014. A primitive placoderm sheds light on the origin of the jawed vertebrate face. Nature 507:500-503.

 

Dupret V., Zhu M., Wang J.Q. 2009. The morphology of Yujiangolepis liujingensis (Placodermi, Arthrodira) from the Pragian of Guangxi (South China) and its phylogenetic significance. Zool. J. Linn. Soc. 157:70-82.

 

Finarelli J.A., Coates M.I. 2011. First tooth-set outside the jaws in a vertebrate. Proceedings of the Royal Society of London B: Biological Sciences:rspb20111107.

 

Finarelli J.A., Coates M.I. 2014. Chondrenchelys problematica (Traquair, 1888) redescribed: a Lower Carboniferous, eel-like holocephalan from Scotland. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 105:35-59.

 

Forey P.L. 1998. History of the Coelacanth Fishes. Chapman and Hall, London.

 

Friedman M. 2007. Styloichthys as the oldest coelacanth: implications for early osteichthyan interrelationships. J. Syst. Palaeontol. 5:289-343.

 

Gagnier P.-Y., Wilson M.V. 1996. Early Devonian acanthodians from northern Canada. Palaeontology 39:241-258.

 

Gagnier P. 1996. Acanthodii. Devonian fishes and plants of Miguasha. Quebec, Canada. Munchen, Freidrich Pfeil Verlag:149-164.

 

Gagnier P., Hanke G., Wilson M. 1999. Tetanopsyrus lindoei gen. et sp. nov., an Early Devonian acanthodian from the Northwest Territories, Canada. Acta Geologica Polonica 49:81-96.

 

Gai Z., Donoghue P.C., Zhu M., Janvier P., Stampanoni M. 2011. Fossil jawless fish from China foreshadows early jawed vertebrate anatomy. Nature 476:324-327.

 

Gardiner B., Bartram A. 1977. The homologies of ventral cranial fissures in osteichthyans. In: Westoll T., Andrews S., Miles R., Walker A., editors. Problems in Vertebrate Evolution. Academic Press, London: p. 227-245.

 

Gardiner B.G. 1984. The relationships of the paleoniscid fishes. a review based on new specimens of Mimia and Moythomasia from the upper Devonian of Western Australia. Bull. Br. Mus. (Nat. Hist.) Geol. 37:173-248.

 

Gardiner B.G., Miles R.S. 1994. Eubrachythoracid arthrodires from Gogo, Western Australia. Zool. J. Linn. Soc. 112:443-477.

 

Giles S., Coates M.I., Garwood R.J., Brazeau M.D., Atwood R., Johanson Z., Friedman M. 2015a. Endoskeletal structure in Cheirolepis (Osteichthyes, Actinopterygii), An early ray-finned fish.

Palaeontology 58:849-870.

 

Giles S., Friedman M. 2014. Virtual reconstruction of endocast anatomy in early ray-finned fishes (Osteichthyes, Actinopterygii). J. Paleontol. 88:636-651.

 

Giles S., Friedman M., Brazeau M.D. 2015b. Osteichthyan-like cranial conditions in an Early Devonian stem gnathostome. Nature 520:82-85.

Giles S., Rücklin M., Donoghue P.C.J. 2013. Histology of “placoderm” dermal skeletons: Implications for the nature of the ancestral gnathostome. J. Morphol. 274:627-644.

 

Goujet D. 1973. Sigaspis, un nouvel arthrodire du Dévonien inférieur du Spitsberg. Palaeontographica Abteilung A 143:73-88.

 

Goujet D. 1975. Dicksonosteus, un nouvel arthrodire du Dévonien du Spitsberg—Remarques sur le squelette viscéral des Dolichothoraci. Colloq. Int. CNRS 218:81-99.

 

Goujet D. 1984. Les poissons placodermes du Spitzberg: Arthrodires Dolichothoraci de la Formation de Wood Bay (Dévonien inférieur). Cahiers de Paléontologie, Section Vertebrés. CNRS, Paris.

 

Grogan E.D., Lund R. 2000. Debeerius ellefseni (Fam. Nov., Gen. Nov., Spec. Nov.), an autodiastylic chondrichthyan from the Mississippian Bear Gulch Limestone of Montana (USA), the relationships of the Chondrichthyes, and comments on gnathostome evolution. J. Morphol. 243:219-245.

 

Gross W. 1935. Histologische studien am aussenskelett fossiler Agnathen und Fische. Palaeontographica Abteilung A 83:1-60.

 

Gross W. 1961. Lunaspis broilii und Lunaspis heroldi aus dem Hunsrückschiefer (Unterdevons, Rheinland). Notizblatt Hessisches Landesamtes für Bodenforschung zu Wiesbaden 89:17-43.

 

Gross W. 1963. Gemuendina stuertzi Traquair. Neuuntersuchung. Sonderdruck aus dem Notizblatt des Hessischen Landesamtes für Bodenforschung zu Wiesbaden 91:36-73.

 

Hanke G.F. 2008. Promesacanthus eppleri n. gen., n. sp., a mesacanthid (Acanthodii, Acanthodiformes) from the Lower Devonian of northern Canada. Geodiversitas 30:287-302.

 

Hanke G.F., Davis S.P. 2008. Redescription of the acanthodian Gladiobranchus probaton

 

Bernacsek & Dineley, 1977, and comments on diplacanthid relationships. Geodiversitas 30:303-330.

 

Hanke G.F., Davis S.P. 2012. A re-examination of Lupopsyrus pygmaeus Bernacsek & Dineley, 1977 (Pisces, Acanthodii). Geodiversitas 34:469-487.

 

Hanke G.F., Davis S.P., Wilson M.V. 2001. New species of the acanthodian genus Tetanopsyrus from northern Canada, and comments on related taxa. J. Vertebr. Paleontol. 21:740-753.

 

Hanke G.F., Wilson M.V. 2006. Anatomy of the Early Devonian acanthodian Brochoadmones milesi based on nearly complete body fossils, with comments on the evolution and development of paired fins. J. Vertebr. Paleontol. 26:526-537.

 

Hanke G.F., Wilson M.V.H. 2004. New teleostome fishes and acanthodian systematics. In: Arratia G., Wilson M.V.H., Cloutier R., editors. Recent advances in the origin and early radiation of vertebrates. Verlag Dr. Friedrich Pfeil, München: p. 189-216.

 

Hanke G.F., Wilson M.V.H. 2010. The putative stem-group chondrichthyans Kathemacanthus and Seretolepis from the Lower Devonian MOTH locality, Mackenzie Mountains, Canada. In: Elliott D.K., Maisey J.G., Yu X., Miao D., editors. Morphology, Phylogeny, and Paleobiogeography of Fossil Fishes, Honoring Meemann Chang. Verlag Dr. Friedrich Pfeil, München: p. 159-182.

 

Harris J.E. 1938. The dorsal spine of Cladoselache. Scientific Publications of the Cleveland Museum of Natural History 8:1-6.

 

Hemmings S.K. 1978. The old red sandstone antiarchs of Scotland: Pterichthyodes and Microbrachius. Palaeontographical Society Monographs 131.

 

Holland T. 2013. Pectoral girdle and fin anatomy of Gogonasus andrewsae Long, 1985: implications for tetrapodomorph limb evolution. J. Morphol. 274:147-164.

 

Holland T. 2014. The endocranial anatomy of Gogonasus andrewsae Long, 1985 revealed through micro CT-scanning. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 105:9-34.

 

Janvier P. 1981. Norselaspis glacialis ng, n. sp. et les relations phylogénétiques entre les Kiaeraspidiens (Osteostraci) du Dévonien inférieur du Spitsberg. Palaeovertebrata 11:19-131.

 

Janvier P. 1985a. Les Céphalaspides du Spitsberg. Anatomie, phylogénie et systématique des Ostéostracés siluro-dévoniens. Révision des Ostéostracés de la formation de Wood Bay (Dévonien inférieur du Spitsberg). Cahiers de Paléontologie, Section Vertebrés. CNRS, Paris.

 

Janvier P. 1985b. Les Thyestidiens (Osteostraci) du Silurien de Saaremaa (Estonie). Ann. Paleontol. 71:83-147.

 

Janvier P., Arsenault M., Desbiens S. 2004. Calcified cartilage in the paired fins of the osteostracan Escuminaspis laticeps (Traquair 1880), from the Late Devonian of Miguasha (Québec, Canada), with a consideration of the early evolution of the pectoral fin endoskeleton in vertebrates. J. Vertebr. Paleontol. 24:773-779.

 

Janvier P., Thanh T.D., Phuong T.H. 1993. A new Early Devonian galeaspid from Bac Thai Province, Vietnam. Palaeontology 36:297-297.

 

Jarvik E. 1972. Middle and upper devonian porolepiformes from East Greenland with special reference to Glyptolepis groenlandica N. Sp.: and a discussion on the structure of the head in the porolepiformes. Palaeontogr. Abt. A Palaeozool-Stratigr. 167:180-214.

 

Jarvik E. 1980. Basic structure and evolution of vertebrates. Academic Press, London.

 

Jessen H. 1980. Lower Devonian Porolepiformes from the Canadian Arctic with special reference to Powichthys thorsteinssoni Jessen. Palaeontographica Abteilung A:180-214.

 

Jia L.-T., Zhu M., Zhao W.-J. 2010. A new antiarch fish from the Upper Devonian Zhongning Formation of Ningxia, China. Palaeoworld 19:136-145.

 

Johanson Z. 1997. New Remigolepis (Placodermi; Antiarchi) from Canowindra, New South Wales, Australia. Geol. Mag. 134:813-846.

 

Lebedev O.A. 1995. Morphology of a new osteolepidid fish from Russia. Bulletin du Muséum national d'histoire naturelle. Section C, Sciences de la terre, paléontologie, géologie, minéralogie 17:287-341.

 

Liu Y.-H. 1991. On a new petalichthyid, Eurycaraspis incilis gen. et sp. nov., from the middle Devonian of Zhanyi, Yunnan. In: Chang M.-M., Liu Y.-H., Zhang G.-R., editors. Early Vertebrates and Related Problems of Evolutionary Biology. Science Press, Beijing: p. 139-177.

 

Liu Y. 1965. New Devonian agnathans of Yunnan. Vertebrata palasiatica 9:125-134.

 

Liu Y. 1975. Lower Devonian agnathans of Yunnan and Sichuan. Vertebrata PalAsiatica 13:202-216.

 

Long J. 1983. A new diplacanthoid acanthodian from the Late Devonian of Victoria. Memoirs of the Association of Australasian Palaeontologists 1:51-65.

 

Long J. 1985. A new osteolepidid fish from the Upper Devonian Gogo Formation, western Australia. Rec. West. Aust. Mus. 12:361-377.

 

Long J. 1997a. Ptyctodontid fishes (Vertebrata, Placodermi) from the Late Devonian Gogo Formation, Western Australia, with a revision of the European genus Ctenurella Ørvig, 1960. Geodiversitas 19:515-555.

 

Long J.A. 1988. New palaeoniscoid fishes from the Late Devonian and Early Carboniferous of Victoria. Memoirs of the Association of Australasian Palaeontologists 7:1-64.

 

Long J.A. 1997b. Ptyctodontid fishes (Vertebrata, Placodermi) from the Late Devonian Gogo Formation, Western Australia, with a revision of the European genus Ctenurella Ørvig, 1960. Geodiversitas 19:515-555.

 

Long J.A., Barwick R.E., Campbell K.S.W. 1997. Osteology and functional morphology of the osteolepiform fish Gogonasus andrewsae Long, 1985, from the Upper Devonian Gogo Formation, Western Australia. Rec. West. Aust. Mus. Suppl. 53:1-89.

 

Long J.A., Mark-Kurik E., Johanson Z., Lee M.S., Young G.C., Min Z., Ahlberg P.E., Newman M., Jones R., Den Blaauwen J. 2015. Copulation in antiarch placoderms and the origin of gnathostome internal fertilization. Nature 517:196-199.

 

Long J.A., Mark-Kurik E., Young G.C. 2014. Taxonomic revision of buchanosteoid placoderms (Arthrodira) from the Early Devonian of south-eastern Australia and Arctic Russia. Aust. J. Zool. 62:26-43.

 

Long J.A., Trinajstic K., Young G.C., Senden T. 2008. Live birth in the Devonian period. Nature 453:650-652.

 

Long J.A., Young G.C., Holland T., Senden T.J., Fitzgerald E.M. 2006. An exceptional Devonian fish from Australia sheds light on tetrapod origins. Nature 444:199-202.

 

Lu J., Zhu M., Long J.A., Zhao W., Senden T.J., Jia L., Qiao T. 2012. The earliest known stem-tetrapod from the Lower Devonian of China. Nat. Commun. 3:1160.

 

Maisey J., Miller R., Turner S. 2009. The braincase of the chondrichthyan Doliodus from the Lower Devonian Campbellton formation of New Brunswick, Canada. Acta Zoologica 90:109-122.

 

Maisey J.G. 1989. Hamiltonichthys mapesi, g. & sp. nov.(Chondrichthyes, Elasmobranchii), from the Upper Pennsylvanian of Kansas. Am. Mus. Novit. 2931.

 

Maisey J.G. 2001. A primitive chondrichthyan braincase from the Middle Devonian of Bolivia. In: Ahlberg P., editor. Major events in early vertebrate evolution. Taylor and Francis, London and New York: p. 263-288.

 

Maisey J.G. 2005. Braincase of the Upper Devonian shark Cladodoides wildungensis (Chondrichthyes, Elasmobranchii), with observations on the braincase in early chondrichthyans. Bull. Am. Mus. Nat. Hist. 288:1-103.

 

Maisey J.G. 2007. The braincase in Paleozoic symmoriiform and cladoselachian sharks. Bull. Am. Mus. Nat. Hist. 307:1-122.

 

Miles R.S. 1967. Observations on the ptyctodont fish, Rhamphodopsis Watson. Journal of the Linnean Society of London, Zoology 47:99-120.

 

Miles R.S. 1971. The Holonematidae (placoderm fishes), a review based on new specimens of Holonema from the Upper Devonian of Western Australia. Philosophical Transactions of the Royal Society of London B 263:101-234.

 

Miles R.S. 1973a. Articulated acanthodian fishes from the old red sandstone of England: with a review of the structure and evolution of the acanthodian shoulder-girdle. Bull. Br. Mus. (Nat. Hist.) Geol. 24:113-213.

 

Miles R.S. 1973b. Relationships of acanthodians. In: Greenwood P.H., Miles R.S., Patterson C., editors. Interrelationships of fishes. Zoological Journal of the Linnean Society, London: p. 63-103.

 

Miles R.S., Westoll T.S. 1968. IX.—The Placoderm Fish Coccosteus cuspidatus Miller ex Agassiz from the Middle Old Red Sandstone of Scotland. Part I. Descriptive Morphology. Trans. R. Soc. Edinb. 67:373-476.

 

Miles R.S., Young G.C. 1977. Placoderm interrelationships reconsidered in the light of new ptyctodontids from Gogo, Western Australia. Linn. Soc. Symp. Ser. 4:123-198.

 

Miller R.F., Cloutier R., Turner S. 2003. The oldest articulated chondrichthyan from the Early Devonian period. Nature 425:501-504.

 

Moy-Thomas J.A. 1935. 26. The Structure and Affinities of Chondrenchelys problematica Tr. Proc. Zool. Soc. Lond., Wiley Online Library, p. 391-404.

 

Moy-Thomas J.A. 1936. On the structure and affinities of the Carboniferous cochliodont Helodus simplex. Geol. Mag. 73:488-503.

 

Newman M.J., Burrow C.J., Den Blaauwen J.L., Davidson R.G. 2014. The Early Devonian acanthodian Euthacanthus macnicoli Powrie, 1864 from the Midland Valley of Scotland. Geodiversitas 36:321-348.

 

Ørvig T. 1960. New finds of acanthodians, arthrodires, crossopterygians, ganoids and dipnoans in the upper Middle Devonian calcareous flags (oberer Plattenkalk) of the Bergisch Gladbach-Paffrath Trough. Paläontologische Zeitschrift 34:295-335.

 

Ørvig T. 1975. Description, with special reference to the dermal skeleton, of a new radotinid arthrodire from the Gedinnian of Arctic Canada. Colloq. Int. CNRS 218:43-71.

 

Pan Z., Zhu M., Zhu Y.A., Jia L. 2015. A new petalichthyid placoderm from the Early Devonian of Yunnan, China. C. R. Palevol 14:125-137.

 

Pearson D.M., Westoll T.S. 1979. The Devonian actinopterygian Cheirolepis Agassiz. Trans. R. Soc. Edinb. 70:337-399.

 

Pradel A., Maisey J.G., Tafforeau P., Janvier P. 2009. An enigmatic gnathostome vertebrate skull from the Middle Devonian of Bolivia. Acta Zoologica 90:123-133.

 

Pradel A., Tafforeau P., Maisey J.G., Janvier P. 2011. A new Paleozoic Symmoriiformes (chondrichthyes) from the Late Carboniferous of Kansas (USA) and cladistic analysis of early chondrichthyans. PLoS One 6:e24938.

 

Qiao T., Zhu M. 2010. Cranial morphology of the Silurian sarcopterygian Guiyu oneiros (Gnathostomata: Osteichthyes). Science China Earth Sciences 53:1836-1848.

 

Rayner D.H. 1952. III.—On the Cranial Structure of an Early Palæoniscid, Kentuckia, gen. nov. Trans. R. Soc. Edinb. 62:53-83.

 

Ritchie A. 1967. Ateleaspis tessellata Traquair, a non-cornuate cephalaspid from the Upper Silurian of Scotland. Zoological Journal of the Linnean Society. 47:69–81.

 

Ritchie A. 1973. Wuttagoonaspis gen. nov., an unusual arthrodire from the Devonian of Western New South Wales, Australia. Palaeontogr. Abt. A Palaeozool-Stratigr. 143:58-72.

 

Ritchie A. 1975. Groenlandaspis in Antarctica, Australia and Europe. Nature 254:569-573.

 

Ritchie A. 2005. Cowralepis, a new genus of phyllolepid fish (Pisces, Placodermi) from the late Middle Devonian of New South Wales, Australia. Proc. Linn. Soc. N. S. W. 126:215-259.

 

Robertson G.M. 1938a. The Tremataspidae; Part I. American Journal of Science:172-206.

 

Robertson G.M. 1938b. The Tremataspidae; Part II. American Journal of Science:273-296.

 

Robertson G.M. 1939. An Upper Silurian vertebrate horizon, with description of a new species, Cephalaspis oeselensis. Trans. Kans. Acad. Sci. 42:357-363.

 

Sallan L., Galimberti A.K. 2015. Body-size reduction in vertebrates following the end-Devonian mass extinction. Science 350:812-815.

 

Sansom R.S. 2009. Phylogeny, classification and character polarity of the Osteostraci (Vertebrata). J. Syst. Palaeontol. 7:95-115.

 

Schaeffer B. 1981. The xenacanth shark neurocranium, with comments on elasmobranch monophyly. Bull. Am. Mus. Nat. Hist. 169:3-66.

 

Schultze H.-P. 1968. Palaeoniscoidea-Schuppen aus dem Unterdevon Australiens und Kanadas und aus dem Mitteldevon Spitzbergens. Bull. Br. Mus. (Nat. Hist.) Geol. 16:341-368.

 

Schultze H.-P., Cumbaa S.L. 2001. Dialipina and the characters of basal actinopterygians. In: Ahlberg P., editor. Major Events in Early Vertebrate Evolution. Taylor and Francis, London and New York: p. 315-332.

 

Schultze H.-P., Zidek J. 1982. Ein primitiver acanthodier (Pisces) aus dem Unterdevon Lettlands. Paläontologische Zeitschrift 56:95-105.

 

Stensiö E.A. 1925. On the Head of the macropetalichthyids with certain remarks on the head of the other arthrodires. Field Museum of Natural History Publications 232:89-198.

 

Stensiö E.A. 1932. The Cephalaspids of Great Britain. British Museum (Natural History), London.

 

Stensiö E.A. 1963a. Anatomical studies on the arthrodiran head. Almqvist & Wiksell, Stockholm.

 

Stensiö E.A. 1963b. The Brain and the Cranial Nerves in Fossil, Lower Craniate Vertebrates.

Universitetsforlaget, Oslo.

 

Stensiö E.A. 1969. Elasmobranchiomorphi Placodermata Arthrodires. In: Piveteau J., editor. Traité de paléontologie. Masson, Paris: p. 71-692.

 

Swartz B.A. 2009. Devonian actinopterygian phylogeny and evolution based on a redescription of

Stegotrachelus finlayi. Zool. J. Linn. Soc. 156:750-784.

 

Taverne L. 1997. Osorioichthys marginis, 'Paléonisciforme' du Famennien de Belgique, et la phylogénie des Actinoptérygiens dévoniens (Pisces). . Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre 67:57-78.

 

Thomson K.S. 1965. The endocranium and associated structures in the Middle Devonian rhipidistian fish Osteolepis. Proceedings of the Linnean Society of London 176:181-195.

 

Trinajstic K., Long J.A. 2009. A new genus and species of Ptyctodont (Placodermi) from the Late Devonian Gneudna Formation, Western Australia, and an analysis of Ptyctodont phylogeny. Geol. Mag. 146:743-760.

 

Trinajstic K., Long J.A., Johanson Z., Young G., Senden T. 2012. New morphological information on the ptyctodontid fishes (Placodermi, Ptyctodontida) from Western Australia. J. Vertebr. Paleontol. 32:757-780.

 

Valiukevicius J. 1992. First articulated Poracanthodes from the Lower Devonian of Severnaya Zemlya. In: Mark-Kurik E., editor. Fossil fishes as living animals. Academy of Sciences of Estonia, Tallinn: p. 193-214.

 

Wang N.-Z., Donoghue P.C., Smith M.M., Sansom I.J. 2005. Histology of the galeaspid dermoskeleton and endoskeleton, and the origin and early evolution of the vertebrate cranial endoskeleton. J. Vertebr. Paleontol. 25:745-756.

 

Wängsjö G. 1952. The Downtonian and Devonian vertebrates of Spitsbergen. IX, Morphologic and systematic studies of the Spitsbergen cephalaspids. Nor. Polarinst. Skr. 97:1-611.

 

Watson D.M.S. 1937. The acanthodian fishes. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences:49-146.

 

Westoll T.S. 1936. On the structures of the dermal ethmoid shield of Osteolepis. Geol. Mag. 73:157- 171.

 

White E. 1958. On Cephalaspis lyelli Agassiz. Palaeontology 1:99-105.

 

White E.I., Toombs H.A. 1972. The buchanosteid arthrodires of Australia. British Museum (Natural History), London.

 

Williams M.E. 1998. A new specimen of Tamiobatis vetustus (Chondrichthyes, Ctenacanthoidea) from the late Devonian Cleveland Shale of Ohio. J. Vertebr. Paleontol. 18:251-260.

 

Young G. 1984. Reconstruction of the jaws and braincase in the Devonian placoderm fish Bothriolepis. Palaeontology 27:635-661.

 

Young G. 1986. The relationships of placoderm fishes. Zool. J. Linn. Soc. 88:1-57.

 

Young G.C. 1979. New information on the structure and relationships of Buchanosteus (Placodermi: Euarthrodira) from the Early Devonian of New South Wales. Zool. J. Linn. Soc. 66:309-352.

 

Young G.C. 1980. A new Early Devonian placoderm from New South Wales, Australia, with a discussion of placoderm phylogeny. Palaeontogr. Abt. A Palaeozool-Stratigr. 167:10-76.

 

Yu X. 1998. A new porolepiform-like fish, Psarolepis romeri, gen. et sp. nov.(Sarcopterygii, Osteichthyes) from the Lower Devonian of Yunnan, China. J. Vertebr. Paleontol. 18:261-274.

 

Zhang G.-R., Wang J.-Q., Wang N.-Z. 2001. The structure of pectoral fin and tail of Yunnanolepidoidei, with a discussion of the pectoral fin of chuchinolepids. Vertebrata PalAsiatica 39:9-19.

 

Zhang M.-M., Yu X.-B. 1981. A new crossopterygian, Youngolepis praecursor, gen. et sp. nov., from lower Devonian of East Yunnan, China. Scientia Sinica 24:89-99.

 

Zhang M. 1980. Preliminary note on a Lower Devonian antiarch from Yunnan, China. Vertebrata PalAsiatica 18:179-&.

 

Zhang M. 1982. The braincase of Youngolepis, a Lower Devonian crossopterygian from Yunnan, south-western China. Swedish museum of Natural History, Stockholm.

 

Zhao W., Zhu M., Jia L. 2001. New discovery of galeaspids from Early Devonian of Wenshan, southeastern Yunnan, China. Vertebrata Pal Asiatica 40:97-113.

 

Zhu M. 1991. New information on Diandongpetalichthys (Placodermi: Petalichthyida). In: Chang M.- M., Liu Y.-H., Zhang G.-R., editors. Early Vertebrates and related problems of evolutionary biology. Science Press, Beijing: p. 171-194.

 

Zhu M. 1996. The phylogeny of the Antiarcha (Placodermi, Pisces), with the description of early Devonian antiarchs from Qujing, Yunnan, China. Bulletin du Muséum national d'histoire naturelle. Section C, Sciences de la terre, paléontologie, géologie, minéralogie 18:233-347.

 

Zhu M., Ahlberg P.E. 2004. The origin of the internal nostril of tetrapods. Nature 432:94-97.

 

Zhu M., Gai Z. 2007. Phylogenetic relationships of galeaspids (Agnatha). Frontiers of Biology in China 2:151-169.

 

Zhu M., Wang W., Yu X. 2010. Meemannia eos, a basal sarcopterygian fish from the Lower Devonian of China–expanded description and significance. In: Elliot D., Maisey J., Yu X., Miao D., editors. Morphology, Phylogeny and Paleobiogeography of Fossil Fishes. Verlag Dr. Friedrich Pfeil, München: p. 199-214.

 

Zhu M., Yu X. 2002. A primitive fish close to the common ancestor of tetrapods and lungfish. Nature 418:767-770.

 

Zhu M., Yu X. 2004. Lower jaw character transitions among major sarcopterygian groups—a survey based on new materials from Yunnan, China. In: Arratia G., Wilson M.V.H., Cloutier R., editors. Recent Advances in the Origin and Early Radiation of Vertebrates. Verlag Dr. Friedrich Pfeil, München: p. 271-286.

 

Zhu M., Yu X. 2009. Stem sarcopterygians have primitive polybasal fin articulation. Biology letters 5:372-375.

 

Zhu M., Yu X., Ahlberg P.E. 2001. A primitive sarcopterygian fish with an eyestalk. Nature 410:81-84. Zhu M., Yu X., Ahlberg P.E., Choo B., Lu J., Qiao T., Qu Q., Zhao W., Jia L., Blom H.,

 

Zhu M., Yu X., Ahlberg P.E., Choo B., Lu J., Qiao T., Qu Q., Zhao W., Jia L., Blom H., Zhu Y.-A. 2013. A Silurian placoderm with osteichthyan-like marginal jaw bones. Nature. 502:188–193.

 

Zhu M., Yu X., Choo B., Wang J., Jia L. 2012. An antiarch placoderm shows that pelvic girdles arose at the root of jawed vertebrates. Biology Letters 8:453-456.

 

Zhu M., Yu X., Janvier P. 1999. A primitive fossil fish sheds light on the origin of bony fishes. Nature 397:607-610.

 

Zhu M., Yu X., Wang W., Zhao W., Jia L. 2006. A primitive fish provides key characters bearing on deep osteichthyan phylogeny. Nature 441:77-80.

 

Zhu M., Zhao W., Jia L., Lu J., Qiao T., Qu Q. 2009. The oldest articulated osteichthyan reveals mosaic gnathostome characters. Nature 458:469-474.

 


 [BMD1]Not included in the reference list. Ugh. FFS.